Introduction part of the movie "The Devil Wears Prada" Andrea Sax who is from Ohio state and graduated North Western university applies to a publishing company in NY to make her dream that be journalist come true. But, the department which she got a interview was the department which published the fashion magazine named “RUWAY” and this department was ruled by Miranda Priestley who was told that she was the most strict person in fashion industry. Miranda’s first assistant, Emily Charlton tries to send Andrea back because Andrea wears unfashionable cloth and she looks that she isn’t interested in fashion at all. At that time, Miranda appears in the office. Suddenly, the office becomes busy. While Andrea was distracted by some worker’s moving such as changing shoes from casual one to high heeled one in quickly and checking lipstick, Miranda who is impeccable and has gorgeous hair and cloth showed up. Although Miranda rejected Andrea in the interview once because of Andrea’s attitude that she didn’t care about her fashion, Miranda reconsidered and decided to hire Andrea because Miranda thought that Andrea might be different from previous assistants who had been dreaming of fashion industry and were useless. It’s the begging of the story that one bumpkin girl who wanted to be a journalist grows gradually under the cruel chief editor for RUNWAY which is called the place to make a trend while she has been being demanded unreasonable orders and being scoled and being knocked about. オハイオ州出身で、ノースウェスタン大学卒業のアンドレア・サックスは、NYで報道記者になる事を夢見てとある出版社に応募します。 しかし彼女が面接へと通されたのは、業界一厳しいと噂のミランダ・プリーストリーが編集長を務める雑誌RUNWAYでした。 ファッションには全く興味がなく、野暮ったいその服装を見て第一アシスタントのエミリーチャールトンは彼女を追い返そうとしますが、そこへミランダが出社してきます。 急に慌ただしくなる社内。 急いで靴をヒールに履き替えたり口紅のチェックをする社員たちの動きにアンドレアが戸惑っていると、ゴージャス服装に髪型、一部の隙のないミランダが現れたのです。 アンドレアを一瞥したミランダは彼女の見た目にこだわらない姿勢に一度は面接をおとしますが、いつもファッション業界を夢見て応募してきては使えなかったアシスタントたちとは少し毛色が違うだろう、と名門大出のアンドレアの採用を決めたのでした。 報道記者になりたかった田舎娘が、流行を作る場所とも言われるRUNWAYの鬼編集長のもとで、無理難題を突きつけられ叱られ揉まれながら煌びやかな世界で成長していく物語の始まりです。